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Intergeneric hybrids of Tripsacum dactyloides and Zea mays were first produced and studied by Mangelsdorf and Reeves (1931)0. Since only unreduced female gametes of these fist generation hybrids functioned, the first backcross to Zea gave 38 chromosome plants of the constitution ZZT, containing two Zea sets each with 10 chromosomes and one Tripsacum set with 18 chromosomes (Mangelsdorf and Reeves 1932). At meiosis in these 38 chromosome plants they reported apparentlyu regular synapsis of the two sets of Zea chromosomes and only occasional synaptic association of Tripsacum and Zea chromosomes, so that gametes having a complete set of Zea chromosomes plus 0 to 18 Tripsacum chromosomes were produced. Since in subsequent backcrosses to Zea as the pollen parent extra Tripsacum chromosomes were transmitted to the progeny with low frequency, Zea plants hyperploid for various numbers of Tripsacum chromosomes were readily isolated for study (Mangelsdorf and Reeves 1939).

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